root › cellular organisms › Eukaryota › Opisthokonta › Fungi › Dikarya › Ascomycota › saccharomyceta › Pezizomycotina › leotiomyceta › Lecanoromycetes › OSLEUM clade › Lecanoromycetidae › Lecanorales › Lecanorineae › Parmeliaceae › Letharia › Letharia columbiana