root › Viruses › Monodnaviria › Shotokuvirae › Cossaviricota › Papovaviricetes › Zurhausenvirales › unclassified Zurhausenvirales › Papillomaviridae › Firstpapillomavirinae › Betapapillomavirus › Betapapillomavirus 1 › Colobus guereza papillomavirus 2